Fitness Variation Due To Sexual Antagonism and Linkage Disequilibrium

Extensive fitness variation for sexually antagonistic characters has been detected in nature. However, current population genetic theory suggests that sexual antagonism is unlikely to play a major role in the maintenance of variation. We present a two-locus model of sexual antagonism that is capable of explaining greater fitness variance at equilibrium than previous single-locus models. The second genetic locus provides additional fitness variance in two complementary ways. First, linked loci can maintain gene variants that are lost in single-locus models of evolution, expanding the opportunity for polymorphism. Second, linkage disequilibrium results between any two sexually antagonistic genes, producing an excess of high and low fitness haplotypes. Our results uncover a unique contribution of conflicting selection pressures to the maintenance of variation, which simpler models that neglect genetic architecture overlook.Organismic and Evolutionary Biolog

Estudio de Viabilidad Económica de una Explotación Agrícola en Almussafes (Valencia)

[ES] El trabajo final de grado se basa en el análisis de inversión de una explotación agrícola situada en el término municipal de Almussafes (Valencia). La inversión se va a centrar en cultivos leñosos, con una vida útil de la inversión de 15 años, y se optará por diferentes especies de cultivos leñosos con objeto de repartir el riesgo de la inversión y planificación de los trabajos en la propia explotación. Se realizada un estudio económico de los cultivos leñosos, para determinar cuáles son los más interesantes desde un punto de vista económico, de gestión y planificación de los trabajos de la explotación. El análisis económico se basará en determinar el VAN, TIR y TIEMPO DE RECUPERACIÓN de la inversión aunque para la elección final se tendrán también en cuenta otros puntos como es la gestión de la mano de obra en la explotación.Orobal Ubeda, D. (2016). Estudio de Viabilidad Económica de una Explotación Agrícola en Almussafes (Valencia). http://hdl.handle.net/10251/65012.TFG

Star Formation Histories of the LEGUS dwarf galaxies. II. Spatially resolved star formation history of the Magellanic irregular NGC 4449

We present a detailed study of the Magellanic irregular galaxy NGC 4449 based on both archival and new photometric data from the Legacy Extragalactic UV Survey, obtained with the Hubble Space Telescope Advanced Camera for Surveys and Wide Field Camera 3. Thanks to its proximity ($D=3.82\pm 0.27$ Mpc) we reach stars 3 magnitudes fainter than the tip of the red giant branch in the F814W filter. The recovered star formation history spans the whole Hubble time, but due to the age-metallicity degeneracy of the red giant branch stars, it is robust only over the lookback time reached by our photometry, i.e. $\sim 3$ Gyr. The most recent peak of star formation is around 10 Myr ago. The average surface density star formation rate over the whole galaxy lifetime is $0.01$ M$_{\odot}$ yr$^{-1}$ kpc$^{-2}$. From our study it emerges that NGC 4449 has experienced a fairly continuous star formation regime in the last 1 Gyr with peaks and dips whose star formation rates differ only by a factor of a few. The very complex and disturbed morphology of NGC 4449 makes it an interesting galaxy for studies of the relationship between interactions and starbursts, and our detailed and spatially resolved analysis of its star formation history does indeed provide some hints on the connection between these two phenomena in this peculiar dwarf galaxy.Comment: 16 pages, 25 figures. Accepted for publication in the Astrophysical Journa

A Tale of 3 Dwarfs: No Extreme Cluster Formation in Extreme Star-Forming Galaxies

Nearly all current simulations predict that outcomes of the star formation process, such as the fraction of stars that form in bound clusters (Gamma), depend on the intensity of star formation activity (SigmaSFR) in the host galaxy. The exact shape and strength of the predicted correlations, however, vary from simulation to simulation. Observational results also remain unclear at this time, because most works have mixed estimates made from very young clusters for galaxies with higher SigmaSFR with those from older clusters for galaxies with lower SigmaSFR. The three blue compact dwarf (BCD) galaxies ESO185-IG13, ESO338-IG04, and Haro11 have played a central role on the observational side because they have some of the highest known SigmaSFR and published values of Gamma. We present new estimates of Gamma for these BCDs in three age intervals (1-10 Myr, 10-100 Myr, 100-400 Myr), based on age-dating which includes Halpha photometry to better discriminate between clusters younger and older than ~10 Myr. We find significantly lower values for Gamma (1-10 Myr) than published previously. The likely reason for the discrepancy is that previous estimates appear to be based on age-reddening results that underestimated ages and overestimated reddening for many clusters, artificially boosting Gamma (1-10 Myr). We also find that fewer stars remain in clusters over time, with ~15-39% in 1-10 Myr, ~5-7% in 10-100 Myr, and ~1-2% in 100-400 Myr clusters. We find no evidence that Gamma increases with SigmaSFR. These results imply that cluster formation efficiency does not vary with star formation intensity in the host galaxy. If confirmed, our results will help guide future assumptions in galaxy-scale simulations of cluster formation and evolution.Comment: Accepted for publication in Ap

Hierarchical Star Formation in Nearby LEGUS Galaxies

Hierarchical structure in ultraviolet images of 12 late-type LEGUS galaxies is studied by determining the numbers and fluxes of nested regions as a function of size from ~1 to ~200 pc, and the number as a function of flux. Two starburst dwarfs, NGC 1705 and NGC 5253, have steeper number-size and flux-size distributions than the others, indicating high fractions of the projected areas filled with star formation. Nine subregions in 7 galaxies have similarly steep number-size slopes, even when the whole galaxies have shallower slopes. The results suggest that hierarchically structured star-forming regions several hundred parsecs or larger represent common unit structures. Small galaxies dominated by only a few of these units tend to be starbursts. The self-similarity of young stellar structures down to parsec scales suggests that star clusters form in the densest parts of a turbulent medium that also forms loose stellar groupings on larger scales. The presence of super star clusters in two of our starburst dwarfs would follow from the observed structure if cloud and stellar subregions more readily coalesce when self-gravity in the unit cell contributes more to the total gravitational potential.Comment: 9 pages, 4 figures, accepted for ApJ